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Show 1876.] MR. A. H. GARROD ON LOPHOTRAGUS MICHIANUS. 761 vertebrae in all. The bones, in the specimen under consideration, especially those of the limbs, are extremely porous and badly marked ; nevertheless, on making a section of the head of the metatarsus, it is apparent that the internal tarsal cuneiform bone has so completely fused with it as to leave no line of demarcation. In the Paris specimens of Elaphodus the tarsus exactly resembles that of Cervulus, and the lateral metacarpals are very nearly lost. In the young female from Moupin the milk canine teeth are in place, their permanent successors appearing, in the dry skull, above them. In the male of the same age from Ningpo, the tusks have a remarkably permanent appearance, and there is no evidence from the condition of the maxillary bones that they belong to the milk series. Such being the case, it must be presumed that the milk canines in the male are shed earlier than in the females, as it is not in accordance with any known facts that they should have persistent pulps which would remove any necessity for their replacement. Anatomy of the Alimentary Canal and other Viscera. The muffle is more considerable than in the Elaphine Deer, but resembles that of the Busina and Muntjacs in extending upwards along the outer border of each nostril as far as its superior margin. The canine tusks protrude an inch below the upper lip, and mark the lower lip at the spots at which they come into contact with them. The palate in front of the intermolar region is transversely ridged by folds of the mucous membrane, slightly crenulated at their free backwardly directed edges. These folds are deficient in the middle line ; and those on one side are not continuous with those of the other, but with the spaces which intervene between them. The intermolar region and the palatal surface behind it are smooth, and black instead of flesh-coloured, as it is anteriorly. The tongue is like that in most ruminating animals, broad near the tip, then narrower, and again slightly broader opposite the intermolar eminence. Its mucous membrane is covered with two kinds of papillae-first the filiform, small, thick-set, short and blunt over the anterior part of the organ, conical and larger in the middle of the intermolar eminence, and secondly the fungiform, disk-shaped and flattened, scattered sparsely over the fore part, and at the sides of the intermolar eminence gradually enlarging and becoming arranged in a linear manner, converging as they run back to form the circumvallate papillae, eleven on one side and twelve on the other. The salivary glands present no special features of interest. The tonsils open each by an orifice situated in the middle of a slight depression. The epiglottis is rounded, with a slight notch in the middle line of its contour. The stomach possesses much the proportions of that of the Musk (Moschus moschiferus)*. In the rumen perhaps the converging left lateral caecal extensions of the upper and lower compartments are slightly longer. The villi are there very close-set, elongated, flat- * Vide P. Z. S. 1875, p. 168. |
