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Show 220 ISLAND LIFE. (PART T. Geographical changes would be still more i~npor~ant, and it is almost impossible to exaggerate the mod1ficatwns of the oraanic world that might result from them. A subsidence of la~d separating a large island from a continent would affect the animals and plants in a variety of ways. It would at once modify the climate, and so produce a series of changes from this cause alone; but more important would be its effect by isolating small groups of individuals of many species and thus altering their relations to the rest of the organic world. Many of these would at once be exterminated, while others, being relieved from competition, might flourish and become modified into new species. Even more striking would be the effects when two continents, or any two land areas which had been long separated, were united by an upheaval of the strait which divided them. Numbers of animals would now be brought into competition for the first time. New enemies and new competitors would appear in every part of the country ; and a struggle would commence which, after many fluctuations, would certainly result in the extinction of some species, the modification of others, and a considerable alteration in the proportionate numbers and the geographical distribution of almost all. Any other changes which led to the intermingling of species whose rang~s were usually separate would produce corresponding results. Thus, increased severity of winter or summer temperature, causing southward migrations and the crowding together of the productions of distinct regions, must inevitably produce a struggle for existence, which would lead to many changes both in the characters and the distribution of animals. Slow elevations of the land would produce another set of changes, by affording an extended area in which the more dominant species might increase their numbers; and, by a greater range and variety of alpjne climates and mountain stations, affording room for the development of new forms of life. Geographical Mutations as a Motive Power in b'ringing about Organic Changes.-Now, if we consider the various geographical changes which, as we have seen, there is good reason to believe have ever been going on in the world, we shall find that the motive power to initiate ·and urae on organic chanaes has never 0 b CHAP. X.] THE RATE OF ORGANIC CHANGE. 221 been wanting. In the first place, every continent, though permanent in a general sense, has been ever subject to innumerable physical and geographical modifications. At one time the total area has increased, and at another has diminished; great plateaus have gradually risen up, and have been eaten out by denudation into mountain and valley; volcanoes have burst forth, and, after accumulating vast masses of eruptive matter, have sunk down beneath the ocean, to be covered up with sedimentary rocks, and at a subsequent period again raised above the surface; and the loci of all these grand revolutions of the earth's surface have changed their position age after age, so that each portion of every continent has again and again been sunk under the ocean waves, formed the bed of some inland sea, or risen high into plateaus and mountain ranges. How great must have been the effects of such changes on every form of organic life! and it is to such as these we may perhaps trace those great changes of the animal world which have seemed to revolutionise it, and have led us to class one geological period as the age of reptiles, another as the age of fishes, and a third as the age of mammals. But such changes as these must necessarily have led to repeated unions and separations of the land masses of the globe, joining together continents which were before divided, and breaking up others into great islands or extensive archipelagoes. Such alterations of the means of transit would probably affect the organic world even more profoundly than the changes of area, of altitude, or of climate, since they afforded the means, at long intervals, of bringing the most diverse forms into competition, and of spreading all the great animal and vegetable types widely over the globe. But the isolation of considerable masses of land for long periods also afforded the means of preservation to many of the lower types, which thus had time to become modified into a variety of distinct forms, some of which became so well adapted to special modes of life that they have continued to exist to the present day, thus affording us examples of the life of early ages which would probably long since have become extinct had they been always subject to the competition of the m.ore highly organised animals. As examples of such excessively |