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Show 282 'l'HE PRINCIPLES OF P.\RT II. have moulted two, three, or four times; and these modifications of plumage are regularly transmitted. Inheritance at Corresponding Seasons of the Year. - 'Vith animals in a state of nature innumera1le instances occur of characters periodically appearing at different seasons. vV e see this with the horns of the stag, and with the fur of arctic animals which becomes thick and white during the winter. Numerous birds acquire bright colours and other decorations during the breeding-season alone. I can throw but little light on this form of inheritance from facts observed under domestication. Pallas states,21 that in Siberia domestic cattle and horses periodically become lighter-coloured during the winter; and I have observed a similar marked change of colour in certain ponies in England. Although I do not know that this tendency to assume a differently coloured coat during different seasons of the year is transmitted, yet it probably is so, as all shades of colour are strongly inherited by the horse. Nor is this form of inheritance, as limited by season, more remarkable than inheritance as limited by age or sex. Inheritance as Limited by Sex.-The equal transmission of characters to both sexes is the commonest form of inheritance, at least with those animals which do not present strongly-marked sexual differences, and indeed with many of these. But characters are not rarely transferred exclusively to that sex, in which they first appeared. Ample evidence on this head has been advanced in my work on Variation under Domestica- ~1 'Novre species Quadrupedum e Glirium 01·dine,' 1778, p. 7. On the transmission of colour by the horse, see 'Variation of Animals, &c. under Domestication,' vol. i. p. 21. Also vol. ii. p. 71, for a general discussion on Inheritance as limited by Sex. CHAP. VIII. SEXUAL SELECTION. 283 tion; but a few instances may here be given. There are breeds of the sheep and goat, in which the horns of the male differ greatly in shape from those of the female ; and these differences, acquired under domestication, are regularly transmitted to the same sex. vVith tortoise-shell cats the females alone, as a general rule, are thus coloured, the males being rusty-red. "\Vith most breeds of the fowl, the characters proper to each sex are transmitted to the same sex alone. So general is this form of transmission that it is an anomaly when we see in certain breeds variations transmitted equally to Loth sexes. There are also certain sub-breeds of the fowl in which the males can hardly be distinO'uished .from each other, whilst the females differ co~siderably in colour. V\Tith the pigeon the sexes of the parent-species do not differ in any external character; nevertheless in certain domesticated breeds the male is differently coloured from the female.22 The wattle in the English Carrier pigeon and the crop in the Pouter are more highly developed in the male than in the female ; and although these characters have been gained through long-continued selection by man, the difference between the two sexes is wholly due to the form of inheritance which bas prevailed ; for it has arisen, not from, but rather in opposition to, the wishes of the breeder. Most of our domestic races have been formed by the accumulation of many slight variations ; and as some of the successive steps have been transmitted to one sex alone, and some to both sexes, we find in the different breeds of the same species all gradations between great sexual dissimilarity and complete similarity. In- ~2 Dr. Chapuis, 'Le Pigeon Voyageur Belge,' 1865, p. 87. Boitard et Corbie, 'Les Pigeons de Volit!re,' &c., 1824, p. 173. |