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Show 324 ALDROV ANDA VESICULOSA. CHAP. XIV. pron1inence. These processes are formed of very delicate membrane lined with a layer of protoplasm; and they sometimes contain aggregated globules of hyaline matter. . Two of the slightly diverging arms are directed towards the circumference, and two towards. the midrib, forming together a sort of Greek cross. Occasionally two of the arms are replaced by one, and then the projection is trifid. We shall see in a future chapter that these projections curiously resemble those found within the bladders of Utricularia more especially of Utricularia montana, although this' genus is not related to Aldrovanda. A narrow rim of the broad fiat exterior part of each lobe is turned inwards, so that, when the lobes are c1os.ed, the exterior surfaces of the in-folded portions come into contact. 1,he edge itself bears a row of conical, flattened, transparent points with broad bases, like the prickles on the stem of a bramble or Rubus. As the rim is infolded; these points are directed towards the midrib, and they appear at first as if they were adapted to prevent the escape of prey; but this can hardly be their chief function, for they are composed of very delicate and highly flexible membrane, which can be easily bent or quite doubled back without being cracked. Nevertheless, the infolded rims, together with the points, must somewhat interfere with the retrograde movement of any small creature, as soon as the lobes begin to close. The circumferential part of the leaf of Aldrovanda thus differs greatly from that of Dionroa; .nor can the points on the rim be considered as. homologous with the spikes round the leaves of Dionroa, as these latter are prolongations of the blade, and not mere epidermic productions. They appear also to serve for a widely different purpose. CHAP. )(IV. ALDROV ANDA VESICULOSA. 325 On the concave gland-bearing portion of the lobes:,. and especially on the midrib, there are numerous, long, finely pointed hairs, which, as Prof. Cohn re~ marks, there can be little doubt are sensitive to a touch, and, when touched, cause the leaf to close. They are formed of two rows of cells, or, according to Cohn, sometimes of four, and do not include any vascular tissue. They differ also from the six sensitive filaments of Dionroa in being colourless, and in having a medial as well as a basal articulation. No doubt it is owing to these two articulations that, notwithstanding their length, they escape being broken when the lobes close. The plants which I received during the early part of October from Kew never opened their leaves, though subjected to a high temperature. After examining the structure of some of them, I experimented on only two, as I hoped that the plants would grow; and I now regret that I did not sacrifice a greater number. A leaf was cut open along the midrib, and the glands examined under a high power. It was then placed in a few drops of an infusion of raw meat. After 3 hrs. 20 m. there was no change, but when . next examined after 23 hrs. 20m., the outer cells of the glands contained, instead of limpid fluid, spherical masses of a granular substance, showing that matter had been absorbed from the infusion. That these glands secrete a fluid which dissolves or digests animal matter out of the bodies of the creatures which the leaves capture, is also highly probable from the analogy of Dionrea. If we rna y trust to the same analogy, the concave and inner portions of the two lobes probably close together by a slow movement, as soon as the glands have absorbed a slight amount of |