Benefits and costs of defense in a neotropical shrub

Benefits and costs are central to optimality theories of plant defense. Benefit is the gain in fitness to reducing herbivory and cost is the loss in fitness to committing resources to defense. We evaluate the benefits and costs of defense in a neotropical shrub, Psychotria horizontalis. Plants were either exposed to herbivores or protected within a cage of fine mesh in three gardens planted in large light gaps on Barro Colorado Island, Panama.